Annabelle is back on the scene after a horrible accident last season while Julie Montagu struggles with being a working mom now that her husband is busy running their family's Mapperton estate. Marissa sets out to throw a lavish Thanksgiving dinner for her society friends, but as the holiday approaches, Juliet and Caroline Stanbury forge an alliance that could spell the end of Juliet's friendship with Marissa.

After recently returning to town, Caroline Fleming throws an eccentric holiday party that entrenches her in the London social scene. Annabelle helps Julie build her confidence while Caroline Stanbury deals with the difficulties of running her business. With Christmas only days away, Juliet and Marissa meet to decide the fate of their damaged friendship.

Much to the chagrin of Julie, Caroline Stanbury mentors her on the emerging health food business. The ladies celebrate New Year's Eve in the penthouse of an exclusive London hotel overlooking the stunning festivities. But the party is cut short when Caroline Stanbury's hilariously over-the-top actions threaten to push Juliet and Julie over the edge...all while wearing unicorn pajamas.

The Ladies navigate the fallout of Caroline Stanbury's New Year's Eve naughtiness. While Caroline and Juliet remain at odds over the fracas, Marissa's fortunes rise with some great press for her Top Dog restaurant. Caroline Fleming visits her Grandmother in Denmark. Caroline Stanbury must put aside the gossip and fight for the doors to remain open at her Gift Library.

Juliette and Annabelle are shocked when Julie extends an olive branch to Caroline Stanbury, despite their fall out from New Year's Eve. Meanwhile, Julie seeks funding for her healthy snack food start-up, and Caroline Stanbury must ask her own investors to keep the Gift Library alive. Marissa seeks Annabelle's stylish advice in anticipation of her new restaurant's launch.

After being on the outs with the ladies for weeks, Juliet attempts to turn a new leaf when she invites everyone out for a night of bowling--but whether she can convince them she's changed remains to be seen. Caroline Stanbury faces the unsettling reality of having to scale down her staff in order to appease investors and keep her business afloat. Julie takes Annabelle and Caroline Fleming to her uncle's Beaulieu estate to generate ideas for saving Mapperton.

After insulting Caroline Fleming for dating a younger man, Marissa tries to salvage their friendship. Julie once again fulfills her reputation as Loose Lips Montagu. Caroline Stanbury struggles to remain emotionally stable while her Gift Library continues its downward spiral.

Marissa and Juliet come back together to watch Annabelle take to the catwalk with Naomi Campbell. Caroline Stanbury reaches a tipping point with Gift Library. Caroline Fleming invites the ladies to her homeland of Demark. The mounting tension between the ladies cannot be contained, as Annabelle voices her opinions-- loudly.

The tension between Annabelle and Caroline Stanbury boils over at Caroline Fleming's castle. The baroness visits her mother's grave and faces her father for the first time since Marissa's meddling. The ladies uncover the reason behind Annabelle's attitude.

In the season finale, Caroline Stanbury adjusts to life without Gift Library. The ladies wonder if Annabelle will trust them enough to show at their final winter bash, where a new Queen Bee is buzzing. Each of the ladies' future is gleamed.